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请教这句话里哪个是谓语动词~?

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请教这句话里哪个是谓语动词~?
Then night fell in the midst of a heavy silence, sometime broken by the cries of the pelican and other night-birds, and the noise of the waves breaking upon the shore, chafing against the rocks, or the panting of some far-off steamer beating the water of the Gulf with its noisy paddles.
这句话的谓语动词和主语是什么啊~~~求解答
请教这句话里哪个是谓语动词~?
整句话的主语是night,谓语是fell.其他动词(有各种形态)都是起修饰作用.