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一道sat语法题求解释~

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一道sat语法题求解释~
Between A.D.350 (and A) 550,Kolomoki,(in what B) is now Georgia,(was one of C) the largest and (most densely D) populated towns north of pre-Columbian Mexico.No error(E)
Kolomoki,(in what B) is now Georgia这句话完整还原是什么?
一道sat语法题求解释~
Between A.D. 350 (and A) 550, Kolomoki, (in what B) is now Georgia, (was one of C) the largest and (most densely D) populated towns north of pre-Columbian Mexico. No error(E)
本句正确
in what is now Georgia 是介词短语 作非限制性定语
其中what is now Georgia是名词性从句 相当于一个名词 作介词in的宾语 是宾语从句
意思也就是Kolomoki was in the place which we now call Georgia
如果用定语从句 应该是 where now Georgia is或in which now Georgia is或 which now Georgia is in
不过意思与原句有出入
也就是now Georgia is in the place which used to be Kolomoki